one of the long, slender feathers on the neck or saddle of certain birds, as the domestic rooster, much used in making artificial flies for anglers.

the neck plumage of a male bird, as the domestic rooster.

hackles,

the erectile hair on the back of an animal's neck

At the sound of footsteps, the dog raised her hackles.

anger, esp. when aroused in a challenging or challenged manner

with one's hackles up.

Angling .

the legs of an artificial fly made with feathers from the neck or saddle of a rooster or other such bird.

hackle fly.

a comb for dressing flax or hemp.

Angling . to equip with a hackle.

to comb, as flax or hemp. ?

raise one's hackles

to arouse one's anger

Such officiousness always raises my hackles.